The 420 MHz band is used in place of the UHF T Band surrounding three Canadian border cities (Buffalo, Cleveland and Detroit).
Refer to the following per NTIA (
http://www.ntia.doc.gov/osmhome/alloctbl/alloctbl.html):
US230--Non-government land mobile service is allocated on a primary basis in the bands 422.1875-425.4875 and 427.1875-429.9875 MHz within 80 kilometers (50 statute miles) of Detroit, MI, and Cleveland, OH, and in the bands 423.8125-425.4875 and 428.8125-429.9875 MHz within 80 kilometers (50 statute miles) of Buffalo, NY.
Essentially Canadian broadcast TV stations are using frequencies within the UHF T Band (470-512 MHz). The 420 MHz spectrum is what was selected as the next best option for these higher population areas.