The Sequoia NF had an extensive microwave network. I heard through the FS radio tech grapevine that they found microwave to be very expensive. They leased some circuits from the state or private industry and had some of their own. At FS owned sites the power bills were quite high. My understanding is that some circuits were replaced with UHF.
The Humboldt-Toiyabe NF has a microwave backbone that it part FS owned and maintained and part State of Nevada owned and maintained. For those repeater sites not linked by microwave UHF links have been installed. Most microwave systems do not link to repeaters on UHF, rather the microwave sites operate as remote bases and just receive and transmit to repeaters like any other base station. Why they chose to use UHF links between the two is unknown to me. I was on that forest when the new system was being built in the late 1980's and helped with the planning of it but never thought to ask why UHF links were being included. My role involved site planning/management and not radio nuts and bolts.
The Humboldt-Toiyabe has no choice but to use microwave. Now that these forests are merged, with the Forest Supervisor's office in Sparks, a comm system covering most of the state is required. There isn't one central dispatcher as the forest is split between 5 dispatch centers. However, I do believe that someone from the Forest Supervisor's office can call someone on any ranger district on the radio. It would be difficult to achieve such using UHF.
My impression of the Sierra NF is that they employ very little or no microwave. They might link Mt. Givens to Musick Mtn., which I believe to be their hub site. However, they don't have remote bases at whatever microwave sites they may have. The Plumas, Sequoia and Angeles use a lot of microwave links and field units can use their radios simplex to contact the dispatcher. It makes it difficult to keep track of things when listening some distance away as you can't hear the ground unit traffic.