If your broadcasting on "repeat" you have to pick an associated tone that is connected to the desired repeater tower in order for the repeat mode to work. Each tower covers a certain geographic area and has a set of specific PL tone numbers assigned to it. The FF's carry a map book for areas they don't know showing the repeater coverage statewide. For CAL FIRE, PL tones are assigned with certain purposes as well. For instance, the PL tone 103.5 is the CAL FIRE designated PL tone for all command frequencies. (A little different for the US Forest Service, they put all of a given National Forest's frequencies on the same tone, so different tones used for the Angeles NF than say the Sierra NF).
Bac to CAL FIRE; If your using "direct", only the engine or crew on the nearby piece of line will hear you since the transmission does not connect with a mountaintop repeater. In both repeat and direct more, the receive (RX) frequency is the same. For instance CAL FIRE Command 1, RX is 151.355 in both the receive and direct mode. The difference is for "repeat" mode the receive frequency (RX) will be paired with a transmit (TX) frequency that is different, that unlocks the desired repeater tower when you push the mic key by broadcasting a sub audible tone. In the case of CAL FIRE Command 1 Repeat: TX 159.300, RX 151.355 Tone 103.5. But In the transmit mode for "direct" both the TX and RX frequency will be the same, example: CAL FIRE 1 Direct: TX 151.355, Receive 151.355...no tone
For scanning purposes, if one was to enter just the RX frequency desired without listing a tone for the frequency and just leave the PL slot open, both repeat and direct mode radio transmissions will be heard since the RX frequency is the same for both. Or, if you live in an area with a lot of interference, you can enter them in your scanner twice, once with a PL tone for repeat, and once without for direct. (The one without the PL tone may be subject to more interference).