The answer is "well it depends". If an incident requires mutual aid at a County border line let's say, the patch can be established and everyone can talk. LASD could use OC VHF,UHF, or 800 access on their APX8000 and OC agencies could stay on Countywide Red tg or even move to an Orange law tac tg. OC Control 1 has the ability to do what ever they need to do. No need for the link patch to be used by OC Control 1 and LASD SCC.
Now if we are talking about a rolling pursuit, that's where "fingertip roaming" occurs. OC units would switch to an 8Call, 8Tac, or CA LAW 8 conventional channel to converse with LA agencies. Their 800 infrastructure is limited but LA RICS is moving towards a multi band mutual aid overlay of the County similar to LARTCS but next gen and will incorporate extensive coverage improvements.
A lot of LA agencies will have, if they do not already, multi band radios in their vehicles. Either standalone RF decks or APX7500/8500 units. They will have all kinds of channels to access. San Diego, Riverside, and San Berdo are already 700 or 800 for their systems and should have a decent amount of 800 (and 700) mutual aid infrastructure in place.
Eventually the So Cal region could become a "System of Systems" using ISSI links and go as far as having coordinated roaming talkgroups among the various P25 systems. The technology is there to make it happen. I wouldn't touch the political aspect with a 50 foot stick though...time will tell.